Guys,
One thing that we should remember is that almost all original rifles had their bores freshened during their life, and probably freshened many times before they arrive at our time. This means that the bore, lands and grooves, were recut. Each freshening process slightly changed the bore configuration - the bore caliber increased by a couple of mils each freshening - start with a 0.442 bore, freshened to a 0.444 bore, then refreshen to a 0.446 and so on. Also the actual final shape of the grooves, width, depth, radius or square, depended more on the history of the gunsmith (or gunsmiths) doing the freshening than on the original barrel maker. The number of grooves and the twist was not changed by freshening, obviously.
Here is a photo of an original rifle bore freshening tool with the cutter in place to freshen the groove. I am sure that you can see that the groove configuration or shape is set by this freshening cutter rather than by the original rifling cutter. It is likely, if not truly provable, that a rifle barrel that started like as a 0.513 round bottom groove has come to us today as a 0.524 square bottom groove. This cutter was used by the Fry brothers in Ligonier Pa in the mid to late 19th c.
Does this make sense?
Jim